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英语定语从句:All of the flowers now raised here have developed fro

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英语定语从句:All of the flowers now raised here have developed from those____in the forest.
A once they grew B they grew once C they once grew D once grew
为什么选c?为什么说省略了宾语which?
1 once 这里是副词,一般放在行为动词前,助动词后 如 Tom has once broken his leg.
2 those是先行词 做后面定语从句的修饰对象,而which在定语从句中是关系代词省略,是定语从句中主语they(花的种植者)的受动对象.those引导定语从句时有关系词省略的情况