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143.Dried foods are invaluable to climbers,explorers and sol

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143.Dried foods are invaluable to climbers,explorers and soldiers in battle,_____ little storage space.
A.which have B.which has C.who have D.that has
选C,先行词是指climbers,explorers and soldiers吗?怎么翻译这句话?
是,who代指以上3者
干粮对于那些没多少储藏空间的攀岩者,探险家和战争中的士兵来说是非常宝贵的