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一题英语选择most of the visitors were from Australia ,____lost a t

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一题英语选择
most of the visitors were from Australia ,____lost a total of five people in Friday's flood
A which B where C what D when
为什么选A
老师说“___”为lost的主语 “what"可以做主语吗?
定语从句缺少主语,用which或that.但是非限制定语从句中不能用that故选用which.其他三个为关系副词.在从句句子成分完整时选用.在这里显然成分不完整.首先排除.至于你说的what是用于名词性从句的