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Ellen was a painter of birds and of nature,_____,for some re

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Ellen was a painter of birds and of nature,_____,for some reason,had withdrawn from all human society A.which B.who C.where D.whom
先行词是啥?为啥是这个?
应该是B,主语从句.Ellen是个画鸟和自然的画家,她,因为一些原因,已经退出人类社会了(不想画关于社会的东西了).